Theo James was born in Oxford, Oxfordshire, England, United Kingdom to Jane (née Martin) and Philip Taptiklis. He is the youngest of five children, having two older brothers and two older sisters. He is of one quarter Greek descent, with the rest of his ancestry being English and Scottish. He graduated high school from Aylesbury Grammar School and earned a degree in Philosophy from the University of Nottingham. He then trained at the Bristol Old Vic Theatre School. He started making guest-star appearances on television shows and had minor roles in films. He then starred as Detective Walter Clark in the crime-drama television series Golden Boy (2013). He rose to fame with his breakthrough role as Tobias “Four” Eaton in the film series Divergent (2014), Insurgent (2015) and Allegiant (2016) based on the novels. Some of his other films, include, Underworld Awakening (2012), The Benefactor (2015), Underworld: Blood Wars (2016), London Fields (2018) and Backstabbing for Beginners (2018).